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Server not accepting messages to remote domains, even with Anonymous users allowed (550 5.7.1)

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Hello all

I've got an odd issue here on an Exchange 2010 SP2/RU4-2008R2 server that is not accepting email for remote domains. I've setup the receive connector as follows:

RunspaceId                              : f2300af2-ecef-435f-935a-bf4b6701bee1
AuthMechanism                           : Tls, Integrated, BasicAuth, BasicAuthRequireTLS, ExchangeServer
Banner                                  :
BinaryMimeEnabled                       : True
Bindings                                : {:::25, 0.0.0.0:25}
ChunkingEnabled                         : True
DefaultDomain                           :
DeliveryStatusNotificationEnabled       : True
EightBitMimeEnabled                     : True
BareLinefeedRejectionEnabled            : False
DomainSecureEnabled                     : True
EnhancedStatusCodesEnabled              : True
LongAddressesEnabled                    : False
OrarEnabled                             : False
SuppressXAnonymousTls                   : False
AdvertiseClientSettings                 : False
Fqdn                                    : SERVERNAME.nt.univ-wea.com
Comment                                 :
Enabled                                 : True
ConnectionTimeout                       : 00:10:00
ConnectionInactivityTimeout             : 00:05:00
MessageRateLimit                        : unlimited
MessageRateSource                       : IPAddress
MaxInboundConnection                    : 5000
MaxInboundConnectionPerSource           : unlimited
MaxInboundConnectionPercentagePerSource : 100
MaxHeaderSize                           : 64 KB (65,536 bytes)
MaxHopCount                             : 60
MaxLocalHopCount                        : 8
MaxLogonFailures                        : 3
MaxMessageSize                          : 30 MB (31,457,280 bytes)
MaxProtocolErrors                       : 5
MaxRecipientsPerMessage                 : 5000
PermissionGroups                        : AnonymousUsers, ExchangeUsers, ExchangeServers, ExchangeLegacyServers

PipeliningEnabled                       : True
ProtocolLoggingLevel                    : Verbose
RemoteIPRanges                          : {0.0.0.0-255.255.255.255, ::-ffff:ffff:ffff:ffff:ffff:ffff:ffff:ffff}
RequireEHLODomain                       : False
RequireTLS                              : False
EnableAuthGSSAPI                        : False
ExtendedProtectionPolicy                : None
LiveCredentialEnabled                   : False
TlsDomainCapabilities                   : {}
Server                                  : SERVERNAME
SizeEnabled                             : EnabledWithoutValue
TarpitInterval                          : 00:00:05
MaxAcknowledgementDelay                 : 00:00:30
AdminDisplayName                        :
ExchangeVersion                         : 0.1 (8.0.535.0)
Name                                    : Default SERVERNAME
DistinguishedName                       : CN=Default SERVERNAME,CN=SMTP Receive Connectors,CN=Protocols,CN=SERVERNAME,C
                                          N=Servers,CN=Exchange Administrative Group (FYDIBOHF23SPDLT),CN=Administrativ
                                          e Groups,CN=RGNAME,CN=Microsoft Exchange,CN=Services,
                                          CN=Configuration,DC=nt,DC=DOMAIN,DC=com
Identity                                : SERVERNAME\Default SERVERNAME
Guid                                    : 75ccba6d-4c90-40b8-bcae-6f677f63d418
ObjectCategory                          : DOMAIN.com/Configuration/Schema/ms-Exch-Smtp-Receive-Connector
ObjectClass                             : {top, msExchSmtpReceiveConnector}
WhenChanged                             : 10/10/2012 1:37:45 PM
WhenCreated                             : 9/14/2012 9:50:58 AM
WhenChangedUTC                          : 10/10/2012 6:37:45 PM
WhenCreatedUTC                          : 9/14/2012 2:50:58 PM
OrganizationId                          :
OriginatingServer                       : DC1.nt.univ-wea.com
IsValid                                 : True

I have restarted the HT service, restarted the server, and tried to delete/recreate the connector but nothing works. The 2007 server that is configured with a similar receive connector works fine. When I telnet to the 2007 server on port 25 I can go through the whole process, but when I try on the 2010 server I get a 550 5.7.1 cannot relay error. I've been looking around for the past few hours but everything tells me this connector is properly configured. Any ideas?


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